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They're At A Stage As Tiny Creatures, One Disheartened

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dvdfrwthr | 09:44 Fri 05th Aug 2022 | Crosswords
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I know answer is PUPAE, but where does 'one disheartened' fit?
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Thanks Etch for your link to the answer from 2014!
I only signed up this year.

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They're At A Stage As Tiny Creatures, One Disheartened

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