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13a Third stage of cell division, during which chromosomes move to the poles of the spindle (8)
23a Line, groove or ridge behind which darts players stand when throwing (4)
27a Greek lyric poet noted for his short celebrations of love and wine, born in Teos (8)
32a Any herbaceous perennial plant of the genus Polygala, especially the common European P. vulgaris (8)
38a English folk-rock singer, lead vocalist for Fairport Convention (5,5)
1d Patient heroine of Chaucer's Clerk's Tale, from a story by Boccaccio (8)
3d Body of permeable rock containing sufficient groundwater to supply wells and springs (7)
17d 'Old Grey Donkey' in the Winnie The Pooh stories of AA Milne (6)
19d Muhammad's flight from Mecca to Medina in AD 622, from Arabic meaning Departure (6)
28d Person who makes and serves coffee in a coffee bar, from Italian meaning bartender (7)
30d Dry pulpy residue left after the extraction of juice from sugar cane, used as fuel and for making paper, etc. (7)
33d Differentiated structural and functional unit in an animal or plant that performs a specialised task (5)