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That site just suggests that Yeoman Warders have always done the job.
The Yeoman Warders are descended from the ancient band of warders who, from early in the Tower's history, had the responsibility for guarding the gates and royal prisoners. From the reign of Henry VIII (1509-47), these duties were carried out by the King's Yeomen at the Tower who were entitled to wear the royal livery - a version of which is still worn. The uniform that most visitors see today is the blue undress uniform introduced in 1858 and worn as the Yeoman Warders' normal working dress. The red and gold state dress is only worn for special occasions, ceremonies, church parades and when royalty visits the Tower. Modern Yeoman Warders (of whom there are about 40) are still Extraordinary Members of the Queen's Bodyguard and are all former warrant officers from Her Majesty's Forces with an honourable service record of at least 22 years.
As well as the Yeoman Warders, the Tower is guarded by a military guard detailed for duty from the same regiments that guard Buckingham Palace and St James's Palace.
Unfortunately it could be anything from wardens, moats, ravens, lions and additional towers. The official site of the Tower of London (not the one above, this one: http://www.hrp.org.uk/webcode/tower_home.asp) doesn't give any indication of any changes to the defences in 1789.
If you find out, let us all know!